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20 Jan 2017 - 12:36 AM  UPDATED 25 Jan 2017 - 10:38 AM

The fourth day of the Australian Open was dominated by the biggest upset in recent tennis history.

A capacity crowd at Rod Laver Arena, and an even bigger one huddled around screens across the world, saw the demise of tennis great Novak Djokovic.

The world number two and defending Australian Open champion was defeated in five sets (6-7 7-5 6-2 7-6 6-4) by wildcard Uzbek Denis Istomin.

The intensity of the match was evident from the outset; the first game lasting an incredible 16 minutes.

At the end of the 4 hours and 50 minutes, it was world number 117 Istomin who was the star of the night.

Istomin says he wouldn’t have dreamt he’d beat Djokovic, especially in five sets.

It’s the first time since Wimbledon 2008 Djokovic hasn’t reached a third round of a grand slam.

The Serbian star says he wasn’t expecting to be exiting the tournament this early.

In the men’s other contests, rising German star Alexander Zverev defeated the USA’s Frances Tiafoe in straight sets (6-2 6-3 6-4).

He’ll face Rafael Nadal in the next round, who defeated Marcos Baghdatis in straight sets (6-3 6-1 6-3).

Meanwhile Serena Williams got a step closer to winning her record 23rd Grand Slam title, defeating Czech Lucie Safarova in straight sets (6-3 6-4).

Williams says as she advances through the tournament, she is reminding herself to always be the Serena she has been.

On the Australian front, Daria Gavrilova made it through to the third round for a second straight year.

Russian-born Gavrilova defeated Croatia’s Ana Konjuh in three sets (6-2 1-6 6-4).

Aussie wildcard Andrew Whittington’s tournament ended at the second round when he was beaten by Croatia’s Ivo Karlovic in straight sets (6-4 6-4 6-4).

Karlovic says the 23-year-old Australian is a promising player.

The tournament also ended for Jordan Thompson, who lost to Austria’s Dominic Thiem in four sets (2-6 1-6 7-6 4-6).

The world number eight dominated for most of the match except for in the third set, when the 75-th ranked Australian fought it to a tiebreak.

Day five of the Australian Open will feature defending champion Angelique Kerber who’ll take on Czech Republic’s Kristyna Pliskova, Australian wildcard player Ashleigh Barty will face Germany’s Mona Barthel and Venus Williams will play China’s Ying-Ying Duan.

In the men’s singles, Stan Wawrinka will take on Serbia’s Viktor Troicki, Roger Federer will face Czech Tomas Berdyh, Kei Nishikori will be up against Slovakia’s Lukas Lacko, Andy Murray will play American Sam Querrey, and for the Australians, Bernard Tomic will attempt to beat Briton Dan Evans.