Fijian-born British Army soldier Semesa Rokoduguni is one of three uncapped players included in the England squad named by coach Stuart Lancaster for next month's internationals at Twickenham.
The dashing wing forced his way into a 33-man squad named on Wednesday for the Tests against New Zealand, South Africa, Samoa and Australia, starting with the visit of the world champion All Blacks on November 8, after impressing in the Premiership this season.
England will face the Wallabies at Twickenham on November 29.
Rokoduguni, 27, who forced his way in ahead of Chris Ashton, was joined by two more uncapped players in flanker Calum Clark and lock George Kruis.
Centre Manu Tuilagi was ruled out of all four Tests by club boss Richard Cockerill because of a groin injury.
But while Tuilagi was not included in Wednesday's squad, he was also missing from a list of players not considered due to injury - suggesting England still have hope of him featuring next month.
"We have included Jonathan Joseph as an injury replacement for Manu," Lancaster said in a Rugby Football Union statement.
"It's obviously disappointing to lose Manu but Jonathan has been in great form and deserves this chance.
"Our medics will continue to work closely with Leicester and ensure that Manu is supported in his rehabilitation from his injury."
Five-eighth Danny Cipriani, the one-time 'golden boy' of Red Rose rugby was not included by Lancaster for a series of matches which mark the start of 2015 World Cup hosts England's run-in to next year's global showpiece.
Instead, Lancaster stuck with Owen Farrell and George Ford, as well as the reliable Stephen Myler, as his three options at No.10.
While there has long been a public debate about England captain Chris Robshaw's worth as a Test backrower, Lancaster has never lost faith in his skipper.
"Chris continues to lead from the front and with other players now stepping up into captaincy and senior roles at their clubs, we are building a team of leaders who can take England forward," Lancaster said.
"We have four quality sides coming to Twickenham for this internationals series and we are really looking forward to playing at home in front of our passionate supporters as we continue the build up to the World Cup in 12 months' time."
Lancaster also has the option of adding to his squad following this weekend's round of European club matches.
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